The Scottish Government's economic development strategy is ineffective and a review of its enterprise services must address ongoing policy failures, the Scottish Conservatives have said.

The Tories attacked the SNP's economic record as MSPs debated phase one of the review at Holyrood.

The party's economy spokesman Dean Lockhart said figures on economic output and productivity targets show Scotland is lagging behind the UK as a whole.

But Economy Secretary Keith Brown said the Tories were down-playing the impact of Brexit, and accused UK Government ministers of "absenting" from their role in the Scottish economy.

Recent figures show output from Scotland grew by 0.4% in the second quarter of this year, compared with 0.7% for the UK.

The Scottish Government outlined a series of reforms to Scotland's enterprise and skills services earlier this week aimed at boosting business, including the creation of a Scotland-wide statutory board to co-ordinate the activities of Scottish Enterprise and Highlands and Islands Enterprise, and a new enterprise agency for the south of Scotland.

"The SNP have had their hands on the levers of the economy for almost a decade. You had a review of the enterprise agencies when you first came to power in 2007, and you've now had another end-to-end review," Mr Lockhart said.

"So I think you've had enough time to establish your economic credentials, and the economic data isn't very promising on your side."

He added: "Given these failures in economic and business development policies I call on the SNP to include in phase two of the report a detailed assessment of how it will address these ongoing policy failures and set out specific steps to increase economic growth in Scotland and productivity in the Scottish economy."

Earlier in the debate, Mr Brown said Scotland's economy has "grown modestly" since the start of 2016, and now faces major challenges in the face of the Brexit vote.

"We have to acknowledge that... we have to step up our performance in order to achieve our ambition, and that level of challenge we face has been increased exponentially by the EU referendum result, creating a new context that requires fresh urgency," he said.

"The Tory tactic of saying we are not as good as the rest of the UK seems to me to be a bizarre one.

"There are two governments involved in the economy of Scotland, and to absent yourself from being involved doesn't seem to be a commendation of that approach."

Labour's Richard Leonard urged the Government to put more money, not less, into economic development.

Scottish Enterprise has seen a real terms budget cut of 16% over the period 2008 to 2015, he said, adding that Highlands and Islands Enterprise had had its funding reduced by 22%.

He said: "If we demand, as I believe we must demand, transformative change in our economy and a rebalancing of our economy with a vibrant manufacturing base, then the Scottish Government needs to be bold and ambitious."