Scientists are to use solar-powered transmitters to track four cuckoos as they fly to their wintering grounds in Africa.

The birds, caught at sites in Lochalsh and Skye, are the most northerly specimens to be used in a study which has previously followed a pair of cuckoos to the Congo Basin and back.

Staff with the British Trust for Ornithology (BTO) attached the transmitter packs to two cuckoos caught on the National Trust for Scotland's Kintail Estate, and also to single birds caught at the trust's nearby Balmacara Estate and Forestry Commission Scotland's Kinloch property on Skye.

The BTO's groundbreaking project is in its third year and is aimed at gaining a better understanding of the migration routes and African wintering grounds cuckoos use.

The cuckoo, which produces one of the most familiar background sounds to early summer in Britain, has undergone a decline in southern England, although numbers in Scotland appear to have remained stable.

An important part of this work will be identifying reasons why the north and west have become refuges for them and suggest ways to ensure this remains the case.