The tragic deaths of two top scientists in a horrific car crash have been described an "immense loss" to Scottish science.

Dr Stewart Rhind and Dr Julian Dawson were passengers in a car when it was involved in a smash with another vehicle on the A90 Dundee to Aberdeen road, just after 4pm on Friday.

The scientists, based at the agricultural research centre the James Hutton Institute, in Aberdeen, died at the scene of the accident just north of Fordoun.

Dr Rhind's wife June, and daughters Joanna and Sarah, paid tribute to the 61-year-old and said he left them with a "wealth of wonderful memories".

In a family statement, they said: "Apart from his research work, his passions were his family, fishing, the natural world and bridge."

Dr Rhind, of Westhill, Aberdeenshire, had collaborated with scientific teams in the US and Australia on animal research issues.

Dr Dawson, 42, was a graduate of Aberdeen University and had returned to the city to work for the Hutton Institute three years ago.

His family said: "Julian was a wonderful man. He will be very much missed by his wife, family and friends."

A spokeswoman for the Hutton Institute said: "Both were valued and respected colleagues who will be sorely missed."

Dr Rhind joined the Hill Farming Research Organisation – a predecessor of the Institute – in 1979 and had worked there ever since.

Dr Dawson and Dr Rhind both died from their injuries at the scene.