MORE than half of Scotland’s upland birds, including the curlew and lapwing, have suffered a “significant long-term decline”, new figures reveal.

A report by Scottish Natural Heritage (SNH) Index Of Abundance For Scottish Terrestrial Breeding Birds,shows 10 of the 17 upland species fell in numbers between 1994 and 2016, contributing to a 16 per cent decrease among upland birds over the period.

Breeding waders including curlew (down 62 per cent), golden plover (-43 per cent), dotterel (-60 per cent), black grouse (-53 per cent) and lapwing (-63 per cent) were among among those suffering the most worrying declines.

SNH says major work is under way to help tackle these declines, including extensive peatland restoration and the Working For Waders project.

However, Scotland’s woodland and farmland birds enjoyed contrasting fortunes, with woodland birds increasing by 67 per cent between 1994 and 2016, and farmland birds increasing by 13 per cent.

The biggest winners were woodland specialists such as the great-spotted woodpecker, which increased by 569 per cent from 1994-2016.

The species has expanded across Europe, possibly as a result of increased forests and woodlands becoming more connected.

The chiffchaff showed the biggest increase of all birds, up 818 per cent up since 1994.

Farmland birds such as the goldfinch (195 per cent), and whitethroat (135 per cent) have also grown in numbers.

Simon Foster, SNH’s trends analyst, said: “It’s wonderful to see that woodland and farmland birds are not only holding their own in Scotland, but that many are thriving.

“However, with some upland birds struggling, there are a lot of people and projects working hard to improve conditions for waders – some of which have seen worrying declines. We and many of our partners are hoping to see these birds fare better in the coming years.”

Terrestrial breeding birds are considered a good biodiversity indicator because they respond quickly to variation in habitat, through changes in breeding output, survival or dispersal.

Since most bird species are relatively easy to identify and count they are often used as indicators of environmental change. SNH used “smoothed indices” -- a statistical way of removing year-to-year fluctuations due to weather conditions or sampling variations -- to track the abundance of Scotland’s terrestrial breeding birds.Data for the report was largely collected by volunteers through the BTO/Joint Nature Conservation Committee (JNCC) Breeding Bird Survey.The figures are published in SNH’s Index of Abundance For Scottish Terrestrial Breeding Birds.

Dr Chris Wernham, head of British Trust for Ornithology Scotland, said: “The Scottish uplands cover a vast area and are a key part of the country’s landscape. Monitoring our upland bird populations presents a unique set of challenges – these are remote, often inaccessible areas, many miles from human population centres.

“It is thanks to the effort and dedication of intrepid volunteers that we are able to produce this report, which highlights worrying declines among a suite of upland bird species, contrasting with increases in many woodland birds.

“Through initiatives within the Breeding Bird Survey, such as Upland Rovers, we hope to see continued improvements in coverage, enabling greater precision in our estimates of change, and in time allowing us to report trends for a greater number of species.”