Nicola Sturgeon has branded as "inhumane" the UK Government's failure to guarantee the rights of EU citizens living in Scotland to remain following the Brexit vote.

With little resolution to the political uncertainty caused by the vote to leave the European Union, the Scottish First Minister stepped up calls for David Cameron and those bidding to succeed him to make such a commitment.

She insisted that people from other countries who had come to the UK and made their lives here "shouldn't be used as a negotiating chip in a wider discussion with Europe".

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The SNP leader has already written to the Prime Minister and the five Tory MPs seeking to replace him, calling for such a guarantee.

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But with none being made, she stepped up her efforts after meeting with European consuls in Edinburgh.

Ms Sturgeon said many of the consuls had "reported concerns and anxiety on the part of their nationals living here in Scotland".

She said: "That is understandable, because there has been a huge question mark placed over their right to live here - not immediately but in the future - as a result of the EU referendum."

READ MORE: Nicola Sturgeon seeks guarantees on rights of EU nationals in Scotland

Since the Brexit vote, Ms Sturgeon and her team of ministers have stressed they welcome the contribution residents from other parts of the EU make.

The First Minister said: "There was a welcome for the message the Scottish Government has given, that we consider EU nationals to be welcome here, that we value their contribution and consider Scotland to be their home. That is a message I am very keen we continue to get across."

But she added: "This uncertainty the EU nationals are living with could be ended at a stroke if the UK Government and all of the candidates for prime minister simply said that their right to live here was guaranteed regardless of what happens in the EU negotiations.

"It's regrettable that hasn't happened on the part of everybody and I call again on the UK Government to give that categoric guarantee to people who have built their lives here, who've got jobs here, pay taxes here, raised families here. They shouldn't be used as a negotiating chip in a wider discussion with Europe.

"There is no doubt that any uncertainty that is being felt could be ended if the UK Government simply gave a guarantee that no matter what happens in the negotiations, people who are already living here, have already built lives here, will have the right to stay. That could be done quickly, simply and straightforwardly and I think it is disgraceful that it hasn't been done."

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Scottish and British people living and working in other parts of Europe "have similar anxieties", Ms Sturgeon said, and she added: "I would hope similar guarantees would be forthcoming from other European countries to them."

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She stressed: "Clearly we would be in a stronger position in arguing for that if the UK was giving guarantees to nationals living here.

"People in both directions that have chosen to live in other countries across the European Union are making contributions, they've built their lives, they shouldn't be used as some kind of negotiating chip, and that is the inhumane aspect to all of this."

The First Minister was also holding talks with Scotland's leading business organisations to hear their concerns about the impact of Brexit.

"Clearly there is a period of significant uncertainty and the risk of a significant impact on the economy ahead of us," Ms Sturgeon said.

She again highlighted her "determination to find a way of protecting Scotland's place in the European Union", but also said: "I'm very keen to work with the business community to get a message out there across Scotland, the UK and Europe as a whole that Scotland remains open for business and we are a good place to do business."

CBI Scotland chair Alison McGregor said afterwards that further work is needed at a UK level on the issues facing EU nationals living in the UK.

She said: "We welcome the First Minister's commitment to allow EU nationals to continue living and working here in Scotland, but we recognise that further work is needed at a UK level. They contribute hugely to our economy and their skills are crucial to the success of our businesses, both now and into the future.

"The Scottish Government should be closely involved in the UK Government negotiations with the EU to help get the best deal for Scottish companies, and business stands ready to play its part too. Companies need access to the single market, trade deals and skills.

"Action to progress long-planned infrastructure projects and national priorities like education should also go ahead as planned. This will help to reassure investors and businesses by signalling confidence in the economy.

"Addressing these issues would be a first step towards providing certainty, and send the right message across the world."

The Herald:

Scottish Secretary David Mundell also held separate talks with business leaders in Edinburgh in the wake of the Brexit vote.

Speaking after the meeting, he said: "Today's positive and constructive meeting was just the start of an ongoing dialogue between Scotland's two governments and the business community in Scotland on the issue of EU exit.

"The overwhelming message from this morning is that Scottish businesses want the UK and Scottish governments to work closely together as part of a 'Team UK' to find a constructive way forward. Our aim should be to unite to ensure the best deal for Scotland and the whole UK as we take forward the necessary work following last week's referendum result.

"We need to work more closely together than ever before to secure the best outcome in these negotiations. As the UK Government's Secretary of State for Scotland, my priority is to ensure that we work closely with the Scottish Government in the months and years ahead and to make sure that Scottish voices and Scottish interests are at the heart of the negotiation process."