The troops from Number Seven Company, Coldstream Guards, began their trek to London from Coldstream in the Scottish Borders in heavy snowfall.
The march will raise money for charity and commemorates an episode in English military and political history.
The soldiers are following the footsteps of General George Monck who led an army on the same route during the English Civil War.
Gen Monck led 6,000 men, part of Oliver Cromwell’s New Model Army, on the march to Parliament 350 years ago.
The 425-mile route is expected to take around 25 days to finish and will help to raise money for the Army Benevolent Fund and the Coldstream charity, helping those injured in recent wars.
Before setting off, the regiment had to be formally granted permission to march by their colonel, Brigadier Jonathan Bourne-May.
March organiser Captain Mark Hayhurst said: “We are on our way.
“There is a good foot of snow on the ground but it is very picturesque.
“We have had to be more direct in our approach, rather than going through the country, we are just using the main road.
“Morale is high. We are just starting off. The sun is shining and we are just cracking on with it.
“The main thing for morale will be getting to know each other a lot better. Quite a few of the soldiers will be deployed to Afghanistan in the next few months, so when things get difficult there they can be up to the mark.”
The march will go through Berwick-upon-Tweed, Newcastle, Durham, York, Selby, Doncaster, Worksop, Mansfield, Loughborough, Market Harborough in Leicestershire, Northampton, Milton Keynes, Dunstable, St Albans, Barnet and Woolwich in London.
The soldiers will stay at Territorial Army centres during the march and aim to arrive at their first stop in Berwick-upon-Tweed by 8pm today, near where the regiment was formed in 1650.
They will attend a special service at the parish church, built by the Coldstream Guards in 1650 as they headed north to fight in the Battle of Dunbar.
The regiment, officially known as Monck’s Regiment of Foot, marched from Coldstream to London in 1660 and was based in the precincts of St James’s Palace to repress rioting.
The company aims to reach Tower Hill in London by January 30.
Why are you making commenting on The Herald only available to subscribers?
It should have been a safe space for informed debate, somewhere for readers to discuss issues around the biggest stories of the day, but all too often the below the line comments on most websites have become bogged down by off-topic discussions and abuse.
heraldscotland.com is tackling this problem by allowing only subscribers to comment.
We are doing this to improve the experience for our loyal readers and we believe it will reduce the ability of trolls and troublemakers, who occasionally find their way onto our site, to abuse our journalists and readers. We also hope it will help the comments section fulfil its promise as a part of Scotland's conversation with itself.
We are lucky at The Herald. We are read by an informed, educated readership who can add their knowledge and insights to our stories.
That is invaluable.
We are making the subscriber-only change to support our valued readers, who tell us they don't want the site cluttered up with irrelevant comments, untruths and abuse.
In the past, the journalist’s job was to collect and distribute information to the audience. Technology means that readers can shape a discussion. We look forward to hearing from you on heraldscotland.com
Comments & Moderation
Readers’ comments: You are personally liable for the content of any comments you upload to this website, so please act responsibly. We do not pre-moderate or monitor readers’ comments appearing on our websites, but we do post-moderate in response to complaints we receive or otherwise when a potential problem comes to our attention. You can make a complaint by using the ‘report this post’ link . We may then apply our discretion under the user terms to amend or delete comments.
Post moderation is undertaken full-time 9am-6pm on weekdays, and on a part-time basis outwith those hours.
Read the rules hereComments are closed on this article