MR John Paul, the director of the Beatson Institute for Cancer

Research in Glasgow from 1966 until 1987, died suddenly on Monday. He

was 72.

Born in Wishaw, he graduated in medicine from Glasgow in 1944 and,

after two years' service as a ship's surgeon in the merchant marine,

embarked on a career in medical research.

After gaining his PhD and working for two years in Edinburgh and New

York, he joined the biochemistry department of Glasgow University as

director of its tissue culture laboratory.

His pioneering work there had a far-reaching influence on almost every

avenue of medical and biological research, with researchers coming from

around the world to learn new techniques for use in the fields of cancer

research, haematology, human genetics, and embryo research.

Mr Paul later moved to the Royal Beatson Memorial Hospital to head its

cancer research laboratories. This gave him opportunities not previously

available to develop his ideas.

The department expanded rapidly under his leadership and he later

founded the Beatson Institute for Cancer Research, which established an

international reputation under his guidance.

Mr Paul was at the forefront of scientific innovation for more than 30

years, until his retirement in 1987. At the height of his career, he was

one of the world's thousand most cited contemporary scientists.

He was admired by his colleagues for his breadth of knowledge, not

simply in medicine and biology but in the physical sciences.

He is survived by his wife, two sons, and a daughter.