The Scottish Government must bring forward "real action" to tackle the problems facing general practice (GP), Liberal Democrat leader Willie Rennie has said.

During exchanges at Holyrood's First Minister's Questions, Mr Rennie said the majority of doctors were not satisfied with the government's plan to combat GP shortages.

Doctors in Glasgow this week warned that surgeries could close in the city as a result of a "GP crisis".

"Can the First Minister tell me when she is going to bring real action forward to put an end to this crisis?" Mr Rennie asked.

Nicola Sturgeon said the government's primary care development fund was providing funding for practices, the testing of a digital service, the recruitment of pharmacists, a GP recruitment and retention programme, and a GP leadership programme.

"I hope that Willie Rennie would accept that that is a fairly impressive list of actions," she said.

Mr Rennie responded: "We have heard it all before. And 99% of GPs who knew about the government's plan said it was simply not enough."

Ms Sturgeon countered that the plan had been welcomed by the chair of the BMA's Scottish GP committee.

"We are shaping primary care so that it is up to meeting the challenges of the future," she said.